Language and Cerebral Cortex Column Rice Mountcastle

Nov 20, 2009 12:33 PM

Upper 3 layers (close to cortex) are called the SUPRAGRANULAR LAYERS I-III.

and are functionally distinct from the INFRAGRANUALR LAYERS  V and V!.

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The upper three layers (closer to the cortical surface) are called the supragranular layers (I-III) and are functionally distinct from the lower two layers (V-VI), which are called the infragranular layers. The supragranular layers project out to other cortical columns, both sending and receiving signals, with layer III cells projecting to adjacent columns and layer II cells projecting to more distant parts of the cortex. The infragranular layers (V-VI) receive input from the supragranular cells of adjacent columns, although they do not reciprocate; instead, they send their signals to extracortical structures, such as the thalamus, and motor and sensory centers.

Layer IV, which divides the column into supragranular and infragranular parts, receives its inputs from the thalamus and transmits signals to the rest of the column. The thalamus receives information from all parts of the cortex and subcortical areas. It works together with the cerebral cortex to create a feedback circuit by passing information from the infragranular cells of the cortex to the thalamus and then back to the layer IV cells of the cortex, with integration occuring both on in the thalamic and cortical centers.

Information flow in the cortex is mediated by the action of the supragranular cells, while additional information from the external and internal environment is integrated by the thalamus and arrives in layer IV of the cortical columns.

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Output to actions is from layer V and Layer V1  (I believe layer V is where Pyramidal cells are in motor cortex.

that lead to contrac. of muscle fibers.

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Outputs: to other columns (II for distant, III for nearby) or to extracortical output (V, VI)

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Language and Cerebral Cortex Column Rice Mountcastle

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