Selection
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<i>Selection</i>
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Although the result of selection, whether through evolution or ontogeny, is structural
coupling (because what is selected is always a structure), selection takes place through
the operational confrontations of a composite system in the medium in which it interacts
as a simple unity through the properties of its components. Thus, it is the differential
effectiveness of the actual operation of different structures of different organisms of
the same kind in parallel existence, or of the same organism in different instances of its
individual history, that constitutes the process of selection in living systems.
Accordingly, selection always takes place in a domain orthogonal to (different from) the
domain of existence of that which is selected. It is this feature of the process of
selection that enables an observer to claim that selection takes place through the
functional value of the structures selected, giving with this judgment, a posteriori, the
misleading impression that what takes place in selection is a semantic coupling that
allows for an infinity of structural realizations. In other words, although the
metaphorical description in functional (semantic) terms is useful for referring to the
orthogonal relation between the domains in which the selective interactions take place and
in which the selected structures exist, the result is structural coupling, because the
operational effectiveness of the selected system depends exclusively on the unique
correspondence thus obtained between its structure and the structure of its medium.
Furthermore, it is also this feature of the process of selection that allows for the
diversity of sequential or simultaneous structural couplings that may take place during
evolutionary or ontogenic adaptation. if the organization of a system is homeostatically
maintained invariant, as occurs in autopoietic systems, adaptation is the homeostatic
clamping through behavior (the actual operation of the autopoietic system in its medium)
of the structural coupling of a system (ontogeny) or of a succession of systems
(evolution) to their static or changing medium
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